Adams, C. A., R. T. Bowyer, J. E. Rowell, W. E. Hauer, and J. 

A. Jenks. 2001. Scent marking by male caribou: an experimental test of rubbing behavior. Rangifer 21:21–27.

Andersson, M., and Y. Iwasa. 1996. Sexual selection. Trends   in Ecology & Evolution 11:53–58.

Bebié, N., and A. G. McElligott. 2006. Female aggression in red deer: Does it indicate competition for mates? Mammalian Biology - Zeitschrift für Säugetierkunde 71:347–355.

Body, G., R. B. Weladji, Ø. Holand, and M. Nieminen. 2014. Highly Competitive Reindeer Males Control Female Behavior during the Rut. PloS one 9:e95618.

Bolker, B. M., M. E. Brooks, C. J. Clark, S. W. Geange, J. R. Poulsen, M. H. H. Stevens, and J.-S. S. White. 2009. Generalized linear mixed models: a practical guide for ecology and evolution. Trends in Ecology & Evolution 24:127–135.

Bolker, B., and R. D. C. Team. 2014. bbmle: Tools for general maximum likelihood estimation. http://CRAN.R-project.org/package=bbmle.

Bro-Jørgensen, J. 2011. Intra-and intersexual conflicts and cooperation in the evolution of mating strategies: lessons learnt from ungulates. Evolutionary Biology 38:28–41. Springer.

Bro-Jørgensen, J., and W. M. Pangle. 2010. Male Topi Antelopes Alarm Snort Deceptively to Retain Females for Mating. The American Naturalist 176:E33–E39.

Burnham, K.P., and Anderson, D.R. 1998. Model selection and inference: a practical information-theoretic approach: 60-64.

Carranza, J., F. Alvarez, and T. Redondo. 1990. Territoriality as a mating strategy in red deer. Animal Behaviour 40:79–88. Elsevier.

Clutton-Brock, T. 2007. Sexual Selection in Males and Females. Science 318:1882–1885.

Clutton-Brock, T. 2009. Sexual selection in females. Animal Behaviour 77:3–11.

Clutton-Brock, T. H. 1989. Mammalian mating systems. Proceedings of the Royal Society of London. Series B, Containing papers of a Biological character. Royal Society (Great Britain) 236:339–372.

Clutton-Brock, T. H., and A. C. Vincent. 1991. Sexual selection and the potential reproductive rates of males and females. Nature 351:58–60.

Clutton-Brock, T. H., K. E. McComb, and J. C. Deutsch. 1996. Multiple factors affect the distribution of females in lek-breeding ungulates: a rejoinder to Carbone and Taborsky. Behavioral Ecology 7:373–378. 

Clutton-Brock, T., and K. McAuliffe. 2009. Female Mate Choice in Mammals. The Quarterly review of biology 84:3–27.

Clutton-Brock, T., and K. McComb. 1993. Experimental tests of copying and mate choice in fallow deer (Dama dama). Behavioral Ecology 4:191–193.
 
Cotton, S., J. Small, and A. Pomiankowski. 2006. Sexual Selection and Condition-Dependent Mate Preferences. Current Biology 16:R755–R765.

Darwin, C. 1871. Sexual selection and the descent of man. Murray, London.

de Jong, K., E. Forsgren, H. Sandvik, and T. Amundsen. 2012. Measuring mating competition correctly: available evidence supports operational sex ratio theory. Behavioral Ecology 23:1170–1177.

de Vries, H. 1998. Finding a dominance order most consistent with a linear hierarchy: a new procedure and review. Animal Behaviour 55:827–843.

Denoël, M., M.-P. Hector, and P. Poncin. 2005. Courtship behavior in the Alpine newt Triturus alpestris at two different densities of males. Herpetologica 61:373–379.

Djaković, N., Ø. Holand, A. L. Hovland, K. H. Røed, R. B. Weladji, E. Fjeldstad, and M. Nieminen. 2012. Association patterns and kinship in female reindeer (Rangifer tarandus) during rut. Acta Ethologica 15:165–171 .

Djaković, N., Ø. Holand, A. L. Hovland, R. B. Weladji, K. H. Røed, and M. Nieminen. 2014. Effects of males’ presence on female behaviour during the rut. Ethology Ecology & Evolution 27:148–160.

Eloranta, E., and M. Nieminen. 1986. Calving of the experimental reindeer herd in Kaamanen during 1970 - 85. Rangifer 6:115–121.

Emlen, S. T., and L. W. Oring. 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197:215–223.

Ginsberg, J. R., and E. J. Milner Gulland. 1994. Sex‐Biased Harvesting and Population Dynamics in Ungulates: Implications for Conservation and Sustainable Use. Conservation Biology 8:157–166.

Grant, J. W. A., and P. E. Foam. 2002. Effect of operational sex ratio on female-female versus male–-male competitive aggression. Canadian Journal of Zoology 80:2242–2246.

Head, M. L., A. K. Lindholm, and R. Brooks. 2007. Operational sex ratio and density do not affect directional selection on male sexual ornaments and behavior. Evolution 62:135–144.

Holand, Ø., L. L’Italien, R. B. Weladji, N. Djaković, K. Røed, A. L. Hovland, and M. Nieminen. 2012. Shit happens–a glimpse into males’ mating tactics in a polygynous ungulate-the reindeer. Rangifer 32:65–72.

Holand, Ø., R. B. Weladji, K. H. Røed, H. Gjøstein, J. Kumpula, J. M. Gaillard, M. E. Smith, and M. Nieminen. 2006. Male age structure influences females’ mass change during rut in a polygynous ungulate: the reindeer (Rangifer tarandus). Behavioral Ecology and Sociobiology 59:682–688. 

Hurlbert, S. H. 1984. Pseudoreplication and the Design of Ecological Field Experiments. Ecological Monographs 54:187–211. Ecological Society of America.

Isvaran, K. 2005. Variation in male mating behaviour within ungulate populations: patterns and processes. Current Science 89:1192–1199.

Jirotkul. 1999a. Operational sex ratio influences female preference and male-male competition in guppies. Animal Behaviour 58:8–8.

Jirotkul. 1999b. Population density influences male-male competition in guppies. Animal Behaviour 58:7–7.

Klug, H., C. I. Kontax, S. Annis, and N. Vasudevan. 2008. Operational sex ratio affects nest tending and aggression in male flagfish Jordanella floridae Goode & Bean. Journal of Fish Biology 72:1295–1305.

Kojola, I. 1991. Influence of age on the reproductive effort of male reindeer. Journal of Mammalogy 72:208–210. 

Kvarnemo, C., and I. Ahnesjo. 1996. The dynamics of operational sex ratios and competition for mates. Trends in Ecology & Evolution 11:404–408. 

Lindenfors, P., J. L. Gittleman, and K. E. Jones. 2007. Sexual size dimorphism in mammals. Sex, size and gender roles: evolutionary studies of sexual size dimorphism:16–26. Oxford University Press Inc. New York.

L’Italien, L., R. B. Weladji, Ø. Holand, K. H. Røed, M. Nieminen, and S. D. Côté. 2012. Mating Group Size and Stability in Reindeer Rangifer tarandus: The Effects of Male Characteristics, Sex Ratio and Male Age Structure. Ethology 118:783–792.

Martin, P., and P. P. G. Bateson. 1993. Measuring behaviour: an introductory guide. Cambridge University Press.

McElligott, A. G., M. P. Gammell, H. C. Harty, D. R. Paini, D. T. Murphy, J. T. Walsh, and T. J. Hayden. 2001. Sexual size dimorphism in fallow deer (Dama dama): do larger, heavier males gain greater mating success? Behavioral Ecology and Sociobiology 49:266–272.

Mills, S. C., and J. D. Reynolds. 2003. Operational sex ratio and alternative reproductive behaviours in the European bitterling, Rhodeus sericeus. Behavioral Ecology and Sociobiology 54:98–104. 

Mysterud, A., Ø. Holand, K. H. Røed, H. Gjøstein, J. Kumpula, and M. Nieminen. 2003. Effects of age, density and sex ratio on reproductive effort in male reindeer (Rangifer tarandus). Journal of Zoology 261:341–344.

Noldus Information Technology. 1998. Matman reference manual. Version 1.1 for Windows. Wageningen, The Netherlands.

Pelletier, F., J. T. Hogg, and M. Festa-Bianchet. 2006. Male mating effort in a polygynous ungulate. Behavioral Ecology and Sociobiology 60:645–654.

Pérez-González, J., and J. Carranza. 2011. Female aggregation interacts with population structure to influence the degree of polygyny in red deer. Animal Behaviour 82:957–970. 

R Core Team (2015). R: A language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria. ISBN 3-900051-07-0, URL http://www.R-project.org/. 

Reynolds, J. D. 1996. Animal breeding systems. Trends in Ecology & Evolution 11:68–72.

Ropstad, E. 2000. Reproduction in female reindeer. Animal Reproduction Science 60-61:561–570.

Røed, K. H., Ø. Holand, M. E. Smith, H. Gjøstein, J. Kumpula, and M. Nieminen. 2002. Reproductive success in reindeer males in a herd with varying sex ratio. Molecular Ecology 11:1239–1243.

Sprenger, D., R. Lange, and N. Anthes. 2011. Population density and group size effects on reproductive behavior in a simultaneous hermaphrodite. BMC Evolutionary Biology 11:107.

Strong, J. S. 2015. Investigation of personality in subdominant male reindeer (Rangifer tarandus) and its relation to somatic cost and reproductive success [MSc thesis]. Concordia University, Montreal.

Tennenhouse, E. M., R. B. Weladji, Ø. Holand, and M. Nieminen. 2012. Timing of reproductive effort differs between young and old dominant male reindeer. Annales Zoologici Fennici 49:152–160.

Tennenhouse, E. M., R. B. Weladji, Ø. Holand, K. H. Røed, and M. Nieminen. 2011. Mating group composition influences somatic costs and activity in rutting dominant male reindeer (Rangifer tarandus). Behavioral Ecology and Sociobiology 65:287–295. 
	
Thiele, J., and B. Markussen. 2012. Potential of GLMM in modelling invasive spread. CAB Reviews 7:1–10.

Uccheddu, S., G. Body, R. B. Weladji, Ø. Holand, and M. Nieminen. 2015. Foraging competition in larger groups overrides harassment avoidance benefits in female reindeer (Rangifer tarandus). Oecologia 3:711–718. 

Weir, L. K., J. W. A. Grant, and J. A. Hutchings. 2011. The Influence of Operational Sex Ratio on the Intensity of Competition for Mates. The American Naturalist 177:167–176.

Young, M. J., L. W. Simmons, and J. P. Evans. 2010. Pre- and post-mating sexual selection both favor large males in a rainbowfish. Behavioral Ecology and Sociobiology 64:915–925.